Live TV production, I think. Mostly in the rackmount form-factor. A plethora of "IO breakout boards" is what turns a regular computer into a "video production system" head-unit.
Though also, at least three of the PCI-e cards shown on the slide were for fibre-optic networking. So, presumably, this would be the Mac to get if you're trying to Beowulf the M2-Ultras together for some kind of NUMA-friendly ML model training. Or just for a render farm. Insofar as Apple dogfoods things, I would guess this is what they use them for themselves.