I didn't read the original article, but this write up does not explain the title at all. Evolution works at the genetic level does it not? What is natural selection modifying if it is not the genes?
Honestly reading that book really opened my eyes to how complex this stuff really is. Biology is unbelievably robust and flexible well beyond the basic model of evolution that we are taught takes places at the genetic level.
Or I wonder if this popular write-up article just got it's title wrong?
The title is technically correct, though misleading.
That quibble aside, there are also epigenetics to consider.
Not quite.
It's the part that filters the ideas, not the part that generates them to begin with.
For that you need a source of change.
It doesn't even need to be external; DNA is automutagenic (read up on transposons).
There's also homologous recombination that exchanges the DNA between the copies of the same chromosome inherited from different parents (if this didn't happen, you couldn't inherit genes from multiple grandparents in a single chromosome).
https://en.wikipedia.org/wiki/Homologous_recombination#In_eu...