Previously they would write in something like C/C++ or use the JVM but try to avoid all garbage collection events though very carefully avoiding allocations, etc.
For them, Rust is a gift!
Also: memory fragmentation is only an issue if you don't have a proper memory management strategy (which you absolute need in any non-trivial code base, even in GC'ed languages).
...which unfounded assertion would that be? As far as I'm aware there are no contenders which would even attempt to throw tools like Maya, 3DSMax, Blender, ... from their throne. And all those tools hail from the 90's.
The only exception in recent time which gained some traction might be Figma for 2D design, but even though it runs in browsers the important parts are also written in C++.