From the literature review section of an article by David Mech, a researcher OP mentions (the whole article is worth reading, and available online here):
> As for high-ranking [wolf] animals asserting any practical control over subordinates, the nature of the interaction is highly conditional. For example, with large prey such as adult moose (Alces alces), pack members of all ranks (ages) gather around a carcass and feed simultaneously, with no rank privilege apparent (Mech 1966; Haber 1977); however, if the prey is smaller, like a musk ox calf, dominant animals (breeders) may feed first and control when subordinates feed (Mech 1988; National Geographic 1988).
> Similarly, pups are subordinate to both parents and to older siblings, yet they are fed preferentially by the parents, and even by their older (dominant) siblings (Mech et al. 1999). On the other hand, parents both dominate older offspring and restrict their food intake when food is scarce, feeding pups instead. Thus, the most practical effect of social dominance is to allow the dominant individual the choice of to whom to allot food.
> The only other rank privilege I am aware of in natural situations is that high-ranking pups are more assertive in competing for food deliveries by adults and sometimes accompany adults on foraging trips at an earlier age than do subordinates (Haber 1977).
https://www.wolf.org/wp-content/uploads/2013/09/267alphastat...