Not just nation state actors, but basically anyone in a position to MITM.
Also, you don't have to be a nation state actor to extort a GH employee. Any bad guy can do a "give me this key or I'll hurt your kid". People are being extorted for a lot less.
There are billions of dollars of assets flowing through GH's infrastructure, for the sake of safety (!= security) of Github's employees, nobody should ever have access to key material.