UUCP with qmail and later Postfix was a great way to shove e-mail over these really unreliable connections. The g protocol worked amazingly well over these unreliable connections because it could pick up where it left off, so as long as you could get a few packets through, eventually your (sometimes large) e-mail messages could make it through, even if they had the spend the night trying.
The company kept that architecture well into the 2010s, even when we all had fairly speedy, reliable Internet connections, because it just worked really well. They eventually went to gmail years after I left.