> Yet they were muscular
We should be careful not to talk past each other. In your previous comment you talked about muscle growth and muscle building. I took it to mean strength building, the one people do in the gym. Hence my comment on untrained individuals and the amount of protein required to build muscle. Anyone who would like to improve their strength and hypertrophy should better be careful to eat enough proteins.
No activity in the nature demands as much strength as the weight training activities in the gym. A small amount of protein may suffice for natives of Papua New Guinea.
As a side note, muscularity might mean different things. A lean individual showing muscular definition might be considered muscular, yet it doesn't mean that they have big muscles or that they are strong. In a similar manner, someone who looks big might look big because they have favorable fat distribution in their body. They may not be strong.
> athletes at the top level can still improve their performance eating 60-65 gram of protein on average per day for over a year.
Yes they can. They are probably mid-distance runners, not strength lifters. Not all types of sports demand high protein intake.