What the article gets wrong, I believe, is the ethical dimension. I would make the argument that for an organism having nociceptors and pain responses is not enough to assume capacity for suffering as we know it (which granted is always a big caveat). I could write a program for a simple robot that shows a pain response. That's not enough.
The missing component here is a brain capable of suffering in a meaningful way. Personally, I suspect there is no convenient limit value for this as you go up the ladder of brain complexity. It's just one big muddy gradient. Where a person draws these lines is almost arbitrary: for example, I don't really care about the well-being of a moskito, but starting with spiders I feel increasingly guilty for not helping them when they're in trouble.