Store and forward.
Do remember that email was THE great federated protocol.
The goal of a mail server was to get your email "at least one hop" closer to your destination. And that wasn't an easy task.
Servers came online and went offline. Users logged in and out. Connections came up and went down. IP wasn't the only transit. DNS? Oh, the hosts file? Even higher things--thing DECnet and Janet.
Email was barely functional most days. Your best bet if you weren't an Internet God and weren't able to write your own super complicated sendmail.cf was to know a sysadmin at a node who had an Internet God and ask him if you could forward emails that you couldn't handle to their server.
Unfortunately, greedy people soon jumped in to take advantage of this generosity, resulting in a tragedy of the commons.
John Gilmore used to run an open relay, and I used to get spam from it. He was really stubborn about promoting the freedom of the spammers over the peace and quiet of the poor recipients. He eventually got shut down, still complaining.
Because the people on the 'net were generally not bad actors back in the day, so we would you need to lock things down?