Nature pays their editors, unlike most society journals. But they are a for-profit business, and are clearly just looking for more profit.

The expected lifetime value of any article in Nature Neuroscience for the authors is well above $11,390, as that is probably 1% of the typical grant that will produce the article, and getting published in this venue makes it much easier to continue to get grants funded, to get top PhD students and postdocs, and to get promoted.

To be clear, though, the authors are not paying for this out of pocket. The funding agencies (i.e. the national government that supports the authors' research) is paying for it. Because many funding agencies now require authors to publish open-acess, Nature can more or less ask whatever price they want.