From what I understand, all this started with SecDB at Goldman, which was a the prototype for all these systems but wasn't Python based. The lore is that SecDB was instrumental in Goldman being able to rapidly figure out what their exposure was during the 2008 crisis.
Some of that team, lead by Kirat Singh went on to start Athena at JP Morgan and then Quartz. I met Kirat once, he was considered a bit of a rock star in the bank tech community. He now runs Beacon, which is basically Bank Python as a service.