It makes me suspect there's something else going on here that's not just good old natural selection.
Have they done any tracking to confirm the tuskless elephants are actually being born to tuskless mothers? Or could it be that the parents are actually affecting the way their offspring are born in some lamarckian-esque sense? If there'd be some trick animals can pull to change their offspring in some ways, that ought to be highly effective for long term species survival and should thus be highly selected for.