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But you already pay them $99/year for a dev account... why do you have to pay them for every purchase as well?

$99/y is really nothing compared to the 30% cut when you look at companies like Epic or Supercell.

I’m certain that they could not continue to provide the App Store in its current form without any fee.

Because the 99$/year are for the dev platform and the 15%/30% cut are for using the infrastructure needed for in app purchases.