Copyright ownership is generally separate from any license
Could you provide some source that corroborates this? Because my understanding is that this has no effect. It merely states something that is already true. That’s what I was hinting at with “possibly meaningless”.
Because all the countries who signed that treaty didn't move to the Berne Convention right away, you had to still reserve your copyright or else lose it in those countries.
The last holdout signed the Berne Convention in 2000, so it's no longer an issue.