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disclaimer: I'm a Firefox user.

I'm genuinely interested in why you think Firefox's UI sucks? What could be done better?

IMO it's more customizable than Chrome's which is a big plus for me.

It's visually unpleasant. Boxy UI elements (notably tabs); elements of different sizes in the same container; too much whitespace in some places; generally not enough whitespace in most places (bookmark folders/menus in general/the whole area around the location bar); elements that change size (the location bar); etc. I don't like customizable UIs; it seems to lead to less attention being paid to the default UI. I don't want to invest in configuring a UI; I just want it to work. Themes degrade over time as they're not maintained. Font rendering is worse than Chrome; colors are different from Chrome (I can't claim that one is right, but it's a jarring difference when comparing them, and I'm used to Chrome). Firefox doesn't have a polished UI, and its whole job is to be a UI for other content.

Written from Firefox, so with that in mind:

In full screen mode, minimize the tab, address, and bookmark bars into the top of the screen. And personally, it bugs me how obscure it is to work with bookmarks (yes, I still use them).

It's still my daily driver at work and home but those are two things that always bug me.

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