This is a big F'N deal and I wouldn't be surprised if it cost Sony more than Microsoft's infamous 1 billion dollar write-down with the Xbox 360's Red Ring of Death.
I don't think this will kill PSN or the PS3, but it's going to significantly dent things. I'm curious to see how much media attention this gets and if we'll see a macro shift towards the Xbox 360 and Wii.
If I was Nintendo and particularly evil I would leverage this opportunity to tout the new system and emphasize the cutting-edge online modes with rock solid security. And MS can also talk about their great track record in the online world.