As for GAN bias: once in Artbreeder the skin tone is entirely my choosing. If there is a bias in the depiction it is my own bias. There might be a slight bias towards a white, brow-haired female but is is more of a gravity that can be avoided with a bit of skill. There is also a challenge in hair that is -for lack of a better explanation- hair made with human intervention (braids, tall wigs, cornrows, dreads etc).
AS for your last suggestion: I've already been thinking of making videos that show a transition of skin ton of lightest plausible to darkest plausible. Tiberius for example I have a very compelling version as a pale-skinned white dude or a dark-skinned person who looks indian/pakistani. Done in paintover/colorization in Photoshop --directly on the bust- where proportions match.
In the second version of the poster I actually challenged myself to do at least do a quick depiction of each emperor with dark skin. Version 1 my approach was to assuming an average based on place of birth. This time I experimented with assume-darkest-features possible simply to test my assumptions and question my biases. Although quick and not robust, I think it was of value.