From the article: "We ought to add some nuance. For each incremental increase in the frequency of the beneficial mutation, it’s not technically a death that’s called for, but only the failure of one individual to survive and reproduce. Maybe someone just doesn’t get to have her allotted offspring. The point is only that the parent’s genetic lineage must come to an end. But in nature, that usually means mortality for either the prospective parent or the offspring. A reasonable shorthand is, well, just death."
This is what happens when skimmers comment.