Unsurprisingly one of those is in a city of 19 million people next to a reservoir of coronaviruses that already caused epidemics.
My understanding of Chinese geography is a bit weak - but it is a bit like if a cow disease broke out in Boston near a BS4 lab. There are certainly some cows in Boston a few blocks from the BS4 lab there. Sure - there are cows in Massachusetts - but statistically it would be a bit surprising that a disease in cows would first show up there.
However, there is a large population of bats right outside Wuhan, and you can find research by the WIV cataloguing different viruses found in the same province. For example : https://pubmed.ncbi.nlm.nih.gov/31009304/
The key here is that the bats the virus (likely) came from are not actually thousands of miles from Wuhan, but only a few dozen. It's just that since the WIV is a huge institute they sometimes also do research in bats in other regions too, because they don't have such facilities.
My understanding is that the virus came from horseshoe bats. These largely live in Southern China. Wuhan is not Southern China. All studies of Corona viruses I can find deal with going to southern China (because that's where horseshoe bats live) to collect samples from horsehoe bats: https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6563315/
Do you have a non-paywalled version to the link you provided? It says they looked at bacteria in bats in the abstract but didn't mention anything about which species of bats.
https://en.m.wikipedia.org/wiki/Shortridge%27s_horseshoe_bat https://en.m.wikipedia.org/wiki/Hubei
You can see that Hubei is indeed in the area where horseshoe bats are found.