This is a beautiful write-up. And I don't wish to detract from the author's work. Nature's result side-by-side with that of the Machine. Makes one feel as though we've taken a step backwards from Alan Turing's "The Chemical Basis of Morphogenesis".
Consider the "morphogenetic puzzle" of a bi-valved seashell that shuts with perfect water-tight seal. There is a constraint to this design: survival!