Most of us have toes that make the front of the foot square-ish when weight-bearing (stand barefoot with your weight on your toes to see what I mean). Walk barefoot on the beach and pay attention to how all of your toes stabilize your stride, relieving your ankles, knees, and hips of some of that strain.
But most sneakers are tapered or at best rounded, squeezing the 3rd through 5th toes inward and preventing them from extending properly.