From the executive summary at the beginning:
The loss of oxygen in the ocean can broadly be put down to two overlying causes – eutrophication as a result of nutrient run-off from land and deposition of nitrogen from the burning of fossil fuels, and the heating of ocean waters as a result of climate change, primarily causing a change in ventilation with the overlying atmosphere so that they hold less soluble oxygen, and compounded by reduced ocean mixing and changes in currents and wind patterns. Ocean deoxygenation is but the latest consequence of our activities on the ocean to be recognized. Ocean warming, ocean deoxygenation, and ocean acidification are major ‘stressors’ on marine systems and typically co-occur because they share a common cause.
Then from Section 2.1, Global Evidence of ocean deoxygenation:
+ Open-ocean deoxygenation is resulting mainly from a warming ocean, increased stratification and changing circulation which interact with eutrophication-induced hypoxia (oxygen concentration below ~60 to 120 µmol O2 kg-1) and biological activity in shelf regions.
+ Oxygen loss is closely related to ocean warming and acidification caused by CO2 increase driven by CO2 emissions as well as biogeochemical consequences related to anthropogenic fertilization of the ocean; hence a combined effort investigating the different stressors will be most beneficial to understand future ocean changes.