My understanding of Google is that they're ok with false negatives because they consider false positives to be far far worse.
You'd be surprised how often the answer to 'Should we hire' has more to do with 'He felt like he would be a dick' rather than 'He doesn't know X and Y technology'.
This sometimes results in candidates who can’t code well or engineers who badly overengineer.
That said, some of the best candidates have been from Google. Perhaps getting those best people was worth a certain percentage of what I consider borderline false positives.
Also, their ahole filter is imperfect. Just as with algorithms, you have people gaming the not an ahole filter.