I think the only ecosystem still trying to match the system’s design is iOS, but given how each version is regressing in its usability, I’m not sure that’s going to last.
Fonts also all match in all desktop applications, and I have configured Firefox to ignore CSS fonts and use the same ones as the rest of the system.
I internally laugh every time somebody gets excited about a new dark theme for their application/OS - like they're coming out of a cave into the 19th century, while I'm living in the future.
There's a plasma-integration plugin for Firefox which makes scrollbars in FF match the Qt scrollbars! It also changes the Firefox scrollbars to click-to-warp instead of click-to-pgup/pgdn, to match with the rest of the system (I believe this happens in all GTK apps though).
There is only one inconsistency I have noticed - in Qt apps you can scrollwheel in inactive windows (configurable if desired), but sometime in the last year GTK changed so that this is no longer possible - I need to switch window focus to scroll GTK windows, except for Firefox. Also Electron apps stand out very much and feel unnatural to use - especially the theming and animations in the File/Edit/View menubar.