In the article it's stipulated it's because of gravity. But it's also mentioned in passing that swimmers hold their breath. I wonder, is there any effect in being able to hold your breath for long when it comes to cardio? As in, I can hold my breath for 4:58, does that offer me any advantage?
Or what about training with a bigger lack of oxygen, as in waiting longer between strokes to breathe or holding your breath while running, does it do any good?