This rarely happens. I would argue that creating an automated system to enforce "rules" should be favored over forcing every developer to know the rules and abides by them.
It is much easier to understand the relationship between css and js when the css is just a js variable.
Having built many large applications using the CSS file method, css modules, each widget gets its own file, co-locating css files with the js, I much prefer using `styled-components`. It provides a ton of flexibility, it opens doors for dynamic css by leveraging js (the hacks I've seen, had to write to get css dynamic is astounding), and I don't have to look very far to see how it's being used because the CSS is literally attached to the component I'm using. All I do in VSCode is cmd+click and bam, I can see the CSS for this component. There's no need for naming convention, there's no worry about global css variables getting into your styles, and there's no digging around looking for css that might be influencing your component. The best part for me is: when I delete the component, the CSS is gone as well. There's no thought put into scouring the FS to find lingering CSS that used to be used.
I love `styled-components` and most of the designers I work with also enjoy it.