4chan was the best. It was meme culture before meme culture. In fact, I believe that's where the modern usage of the word comes from (ytmnd used "fad" for the same concept). I spent so much time on there 15 years ago when I first entered college.
The confusion is that the latter meaning of "meme" has started drifting from "an image macro that spreads" to "an image macro".
The later misuse of the word meme to refer only to image macros is also a separate shift.