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I use TiddlyWiki for note taking, and I think it has most everything you’re asking for. Except, my one complaint is that it uses an odd dialect of markdown.

You'll probably like CintaNotes (Windows only) or Zim Wiki.

Biggest downside to both is the lack of mobile apps. But Zim uses plain text files and Cinta can sync with Simplenote. I mention Cinta mostly because it has some of the better search capabilities I have seen in a note app.

You should check out notion.so

Second this. We use notion for all of our company knowledge-tracking and documentation. It's even great for the more technical side of things, as it supports code-formatting as well.

Check out https://www.wikiful.com

Prioritizes speed & simplicity. Has markdown shortcuts.

(I’m CTO / co-founder)