What they did by centralizing the collection was the best thing they could do to save it, but apparently they didn't even have money to protect a single museum from fire hazard. Spread the collection around the country and you don't even have money/people for proper maintenance and protection against theft in each individual location.
In any case, the strategy they chose obviously didn't work and they lost it 100%.
The strategy didn’t work since any strategy was likely deemed for failure due lack of funding, not because it was suboptimal.
If the likelihood of a fire starting in a non-residential building is roughly proportional to the square footage of the building, then you'd have roughly the same total number of fires across all buildings but 1/N of the collection destroyed each time.
That's not often the case. These museums were built to house these collections, and often include rooms full of drawers, compacting shelving etc with the collected objects. This incudes the basement, but not only the basement.
For example, the Natural History Museum in London has "the majority" of their 80 million specimens stored in the museum in London.
You may be able to go on a behind-the-scenes tour of your local museum.