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I am not very familiar with this project but I see it uses UPnP. Does it rely on UPnP or is it only opportunistic use? If the former, what happens if the user's custom router does not run a UPnP daemon, or if the user prefers to disable UPnP in her consumer router?

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It does rely on being able to communicate through that port - UPnP is one of the options it uses to ensure it can receive external requests. If you run using the tor option it evades firewalls via tor.

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