Why do all these features of a colony need to be part of the same thing?

I see 1. Reputation 2. Governance based on reputation 3. Division of labor 4. Team membership 5. Organizational ownership 6. Dispute resolution

It is not obvious what this particular set of organizational features is necessary or sufficient for. Even worse there are likely many different ways of implementing them and it is not obvious how implementation details will affect resulting organizational dynamics. A better approach would be a modular organizational design with different plug-ins that satisfy different organizational requirements. This would allow organizations to choose their rules and to innovate instead of having to pick up a complex model of organization off of a shelf. A modular design would let you start small, build a minimum set of requirements to power some organizations and then scale up the complexity from there.