I hate that line, you know what most of these companies have in common? They where started by guys with only a few years in the field and a vision. Some of them not the best developers in the world, just guys with a great idea for a product and a passion. The other thing they have in common, is that they have spent millions if not more on rolling their own solutions to compensate for a technology decision they made a long time ago in a garage.
This is the way it works, the new guys come in, they say well company X uses this, they build something on it because X is using it and they hit growing pains at a certain point.
Twitter is experiencing those growing pains now, fortunately they seem to be throwing out the bad and trying new things. A good deal of companies become dogmatic about their choice and given their success just throw money at it to build compensating workarounds.
MySQL was not chosen by those companies because of its merits, it was chosen because those companies started in a garage, and the technical selection was "what are the other guys using in their garage".