There's one difference, though: conventional Python is indented four, not two, spaces. In fact, the main aspect of his "normal" C code that makes it more readable (to me, at least) is his use of four spaces for indentation.
Personally I think the biggest barrier to Lisp readability is too little indentation (two spaces is typical). Looking at his reformatted Lisp, the way I identified the nesting level of his highlighted line was not by looking at trailing parens but by looking at the indentation, which was much wider (and therefore clearer) than standard Lisp.