It is not a secret that kernel devs have a love-hate relationship with systemd, and have for years. That's why Linus said, "You all presumably know why." He was speaking with context that everyone on the LKML already knows.
First result when I google it (but it's been a thing for a long time and on many fronts with many interesting bugs and arguments): http://www.phoronix.com/scan.php?page=news_item&px=MTY1MzA
I don't really subscribe to the "systemd is terrible and is the worst thing that ever happened to Linux" theory, and I'm not trying to be one of those folks that takes every opportunity to make a thread about how much systemd sucks. I like systemd. I mean it when I say that. But, in this case, it seems obvious it is about systemd. I have a hard time coming up with another plausible theory.
Edit: Also, now that I'm thinking about Linus and Poettering butting heads...well, that's hilarious. A couple of brilliant assholes arguing is funny to me. I dunno how much it's actually happened (and how much happened through proxies), but still. Funny.