Ah yes, implied nudity. You know, like clothes. ;)
Here's the iPhone dev agreement: http://www.eff.org/files/20100302_iphone_dev_agr.pdf
The only pertinent section I can find is:
"Applications must not contain any obscene, pornographic, offensive or
defamatory content or materials of any kind (text, graphics, images, photographs, etc.), or other content or materials that in Apple's reasonable judgment may be found objectionable by iPhone or iPod touch users."
Which, in matter of fact, does not prohibit nudity, but rather anything Apple thinks its users might be offended by. Nudity is in fact allowed. We can surmise this from the approval of "Fabulous nude paintings puzzle", and dozens more like it.
http://itunes.apple.com/us/app/fabulous-nude-paintings-puzzl...
Stranger still, "iGirl uncut", where you can "Blow her, shake her, touch her and more", must not be obscene, objectionable, or pornographic.
http://itunes.apple.com/us/app/igirl-uncut/id300462216?mt=8
I agree with you: the panels are sexual. But Apple's policy clearly allows them to ban whatever the hell they feel like. When the enforcement of that policy is wildly inconsistent, it looks a hell of a lot like unjust content regulation to me.
As for the article's claim of homophobia, I don't have the grounds to claim the reviewers who censored this app were motivated by anti-gay sentiment, but I also wouldn't be surprised.