"Average life expectancy at birth was 43 years between 1950-89 and increased to about 50 during the period 1990-2002. Half the population is under 15 years of age."

http://www.philosophy.dept.shef.ac.uk/culture&mind/people/gu... (that is the site of one of the anthropologists quoted in the BBC article)

They also have very high parasite load, which was kind of mentioned in the article; may influence fertility as well as heart health and overall longevity: https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3529658/