Here is a link to the full html article [1] and the pdf [2] The links are a bit hard to find from the abstract page. The html version has some messed up formatting but the pdf is fine. The DOI is: 10.1073/pnas.1601040113

[1] http://www.pnas.org/content/113/33/9232.full [2] http://www.pnas.org/content/113/33/9232.full.pdf