Between the ZFS-like features and the advertised "novel copy-on-write metadata" scheme, I would not be surprised if it was partially based on DragonFly's HAMMER/HAMMER2 filesystem.
(That does leave me wondering how interesting an open-source APFS would be without an open-source Core Storage.)
hammer.ko is 494KB in the latest DFBSD, for those wondering the obvious.
It would be fantastic if they actively helped with HAMMER2 development.